Em Earlville, uma cidade do centro-oeste, Irma (Jessica Lange) e Roy Applewood (Tom Wilkinson) estão comemorando 25 anos de casados. Roy revela para Irma e o reverendo Dale Muncie (Randall Arney) que é uma mulher no corpo de um homem e isto está sendo intolerável, assim pretende fazer uma operação para mudança de sexo. Ele passa a usar cada vez mais adereços femininos, com esta atitude sendo vista com intolerância e repúdio pelos colegas de trabalho e membros da sua igreja. Apesar do choque inicial, Irma tenta entendê-lo e apoiá-lo nesta trajetória, que o faz se tornar "Ruth".
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