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Andrea Sachs (Anne Hathaway) é uma jovem que conseguiu um emprego na Runaway Magazine, a mais importante revista de moda de Nova York. Ela passa a trabalhar como assistente de Miranda Priestly (Meryl Streep), principal executiva da revista. Apesar da chance que muitos sonhariam em conseguir, logo Andrea nota que trabalhar com Miranda não é tão simples assim.
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