Depois que as quatro amigas terminaram o colégio, cada uma toma um rumo diferente na vida e já não estão mais sempre juntas como costumavam estar. Apesar das mudanças em suas vidas, elas precisam uma da outra para superar alguns problemas pessoais. Lena sofre com desencontros amorosos, Tibby tem medo de se entregar à felicidade, Bridget passa por conflitos familiares e Carmen passa por mudanças. Quem sofre com o descaso das meninas é a calça da Irmandade, que mais uma vez vai unir as garotas.
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